Why the “four fundamental forces” view is mistaken

Over at Sandwalk, Larry Moran posted some interesting bits rrom his molecular evolution class exam, including a passage from Mike Lynch arguing for his claim that “nothing in evolution makes sense except in the light of population genetics”.  In this passage, which I’ll quote below, Lynch says that evolution is governed by 4 fundamental forces.

The idea that evolution is governed by population-genetic “forces” is common but fundamentally mistaken.  I wish we could just put this to rest.